TOLEDO, Ohio (AP) _ Two of five chimpanzees that got out of their cage at the Toledo Zoo on Sunday were shot, one fatally, after one bit a keeper and the others rushed an emergency team, a zoo official said.

No visitors were in the area at the time of the incident, said Glenous Favata, curator of mammals. There were only few people in the zoo because it had just opened for the day, she said.

One chimpanzee, a 14-year-old female, was killed and another chimp, a 30- year-old female, was seriously wounded as the emergency team tried to corral the five chimps, Ms. Favata said. A third chimp eventually returned to its cage on its own while the two others were tranquilized, she said.

Ms. Favata said the keeper was treated and released at a hospital after suffering a bite on a finger and bruises on her face. Her name was not made public.

The chimps' cage in the Great Ape Building was apparently left unlocked by a cleaning worker, Ms. Favata said.